Quote:one page (f55r) still has the (unfinished) red lead sketch-lines, so that’s probably how all the drawings were done – a master making a (pencil) sketch and then a scribe tracing & finishing them, before rubbing off the red lead.

My guess is that the marking up for the text was similarly done in red lead, which was similarly then removed by the scribe. I don’t know if there’s any literature on red lead markings, though: that might be a scribal practice local to certain areas. Something to think about, anyway.