Part of Zodiac was added later?

[DONJCH]

hi, I can't understand some words.
please teach me their?(´-ω-`)
I don't understand when read VM product(´・ω・`)


1. Quire
2. bifolium
3. Quiremarks

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The way I think of it is this:

A bifolium is a sheet of parchment folded down the middle to make 4 "pages"
A quire is several of these stitched together to make a "chapter"
Quiremarks are the labels on the quires 1 2 3 4 ...

(Others more experienced may dispute this)