Did Voynich manuscript illustrator never see women breasts?

[Rafal]

But it is evident to me that his skill at drawing them improved enormously and gradually as he worked through the Zodiac section.

My feeling is that he first drew a lot of nymphs and then made some corrections to them.

If you look at the nymphs you may notice that in many cases their right breast (right physically and left on the picture) is darker. I suppose they were added later.

In small number of cases he forgot or didn't care to add this breast, however. I marked such a case for you.
   

When you look at a woman exactly from a profile, she has just one breast visible, right?



So our artist could follow this rule but then realized that his nymphs aren't exactly shown from profile and so they need the second breast